I solved the problem. Apparently, the apachectl command for starting the Apache 2.0.63 server (which I installed for Webwork2) is lower in the $PATH than for the built-in Apache 2.2.28 server (which I didn't know about until now).
Fixing the path got the correct server working.
-Tom
Installation
Apache modules & Webwork: 404: File not found error
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